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accumulation acid active transport blue light Calvin cycle caused closure caused stomatal opening cells and subsidiary changes chlorophyll chloroplasts Commelina communis concentration Cyclamen persicum dark rhythm decrease detached epidermis diurnal effect of light epidermal cells epidermal strips evidence factors free air Further Reading guard cells guard-cell starch Heath hydrolysis illumination increased intact leaves intercellular spaces ions isolated stomata leaf leaf discs leaf surface light effect light intensity malate mannitol Mansfield mechanism Meidner mesophyll metabolic Milthorpe Mouravieff nitrogen occur onion Open stoma opening in light osmotic potential Pelargonium phase photophosphorylation photosynthesis phytochrome plants porometer readings red light reduced showed solutions species starch content stomata closed stomatal cells stomatal closure stomatal movement stomatal responses stomatal starch strips taken subsidiary cells substomatal cavities sugar temperature Tradescantia virginiana turgor uptake values indicate wider Vicia faba water stress water vapor wheat wilting Xanthium Xanthium pennsylvanicum